Original URL: http://www.inform.umd.edu/PBIO/pb250/dahl3.html
PBIO 250 Lecture Notes
James L. Reveal
Norton-Brown Herbarium, University of Maryland
Dahlgren System of Angiosperm Classification
19. Asteranae Takht. (1967)
Campanulanae Takht. ex Reveal (1992)
1. Campanulales Rchb.f. (1828)
1. Pentaphragmataceae J. Agardh (1858)
2. Campanulaceae Juss., nom. cons. (1789)
Cyananthaceae J. Agardh (1858)
Cyphiaceae A. DC. (1839)
Cyphocarpaceae (Miers) Reveal & Hoogl. (1996)
Jasionaceae Dumort. (1829)
Nemacladaceae Nutt. (1842)
Pongatiaceae Endl. ex Meisn., nom. illeg. (1839)
Sphenocleaceae Mart. ex DC., nom. cons. (1839)
3. Lobeliaceae R. Br., nom. cons. (1817)
Dortmannaceae Rupr. (1856)
2. Asterales Lindl. (1833)
Ambrosiales Dumort. (1829)
Carduales Small (1903)
1. Asteraceae Dumort., nom. cons. (1822)
Acarnaceae Link (1829)
Ambrosiaceae Link, nom. cons. (1829)
Anthemidaceae Martinov (1820)
Aposeridaceae Raf. (1838)
Arctotaceae Bessey (1914)
Artemisiaceae Martinov (1820)
Athanasiaceae Martinov (1820)
Calendulaceae Link (1829)
Carduaceae Dumort. (1822)
Centaureaceae Martinov (1820)
Cichoriaceae Juss., nom. cons. (1789)
Cnicaceae Vest (1818)
Compositae Giseke, nom. cons. et nom. alt. (1792)
Coreopsidaceae Link (1829)
Cynaraceae Durande (1782)
Echinopaceae Dumort., nom. cons. (1822)
Eupatoriaceae Martinov (1820)
Gnaphaliaceae Link ex Rudolphi (1830)
Heleniaceae Raf. (1824)
Helianthaceae Dumort. (1822)
Helichrysaceae Link (1829)
Inulaceae Bessey (1914)
Lactucaceae Drude (1879)
Lampsanaceae Martinov, nom. illeg. (1820)
Matricariaceae Voigt (1845)
Mutisiaceae Burnett (1835)
Nassauviaceae Burmeist. (1837)
Partheniaceae Link (1829)
Perdiciaceae Link (1829)
Picridaceae Martinov (1820)
Santolinaceae Martinov (1820)
Senecionaceae Spenn. (1834)
Serratulaceae Martinov (1820)
Tanacetaceae Vest (1818)
Vernoniaceae Burmeist. (1837)
Xanthiaceae Vest (1818)
20. Solananae R. Dahlgren ex Reveal (1992)
1. Solanales Dumort. (1829)
Convolvulales Dumort. (1829)
Nolanales Lindl. (1835)
Polemoniales Bromhead (1838)
1. Solanaceae Juss., nom. cons. (1789)
Atropaceae Martinov (1820)
Cestraceae Schltdl. (1833)
Daturaceae Raf. (1828)
Duckeodendraceae Kuhlm. (1950)
Hyoscyamaceae Vest (1818)
Lyciaceae Raf. (1840)
Nicotianaceae Martinov (1820)
Nolanaceae Dumort., nom. cons. (1829)
Salpiglossidaceae (Benth.) Hutch. (1969)
2. Sclerophylacaceae Miers (1848)
3. Goetzeaceae Miers ex Airy Shaw (1964)
4. Convolvulaceae Juss., nom. cons. (1789)
Cressaceae Raf. (1821)
Dichondraceae Dumort., nom. cons. (1829)
Erycibaceae Endl. ex Meisn. (1840)
Humbertiaceae Pichon, nom. cons. (1947)
Poranaceae J. Agardh (1858)
5. Cuscutaceae (Dumort.) Dumort., nom. cons. (1829)
6. Cobaeaceae D. Don (1824)
7. Polemoniaceae Juss., nom. cons. (1789)
2. Boraginales Dumort. (1829)
1. Hydrophyllaceae R. Br., nom. cons. (1817)
Ellisiaceae Bercht. & J. Presl (1820)
Eutocaceae Horan. (1847)
Hydroleaceae Bercht. & J. Presl (1820)
Sagoneaceae Martinov (1820)
2. Ehretiaceae Mart. ex Lindl., nom. cons. (1830)
Cordiaceae R. Br. ex Dumort., nom. cons. (1829)
Sebestenaceae Vent. (1799)
3. Boraginaceae Juss., nom. cons. (1789)
Anchusaceae Vest (1818)
Buglossaceae Hoffmanns. & Link (1809)
Cerinthaceae Martinov (1820)
Echiaceae Raf. (1837)
Heliotropiaceae Schrad. (1819)
Onosmaceae Martinov (1820)
Wellstediaceae (Pilg.) Novák (1943)
4. Lennoaceae Solms, nom. cons. (1870)
5. Hoplestigmataceae Engl. & Gilg, nom. cons. (1924)
21. Ericanae Takht. (1967)
Sarracenianae Thorne ex Reveal (1996)
1. Bruniales Dumort. (1829)
1. Bruniaceae R. Br. ex DC., nom. cons. (1825)
2. Grubbiaceae Endl., nom. cons. (1839)
Berzeliaceae Nakai (1943)
Ophiraceae Arn. (1841)
2. Fouquieriales Takht. ex Reveal (1992)
1. Fouquieriaceae DC. (1828)
3. Ericales Dumort. (1829)
Actinidiales Takht. ex Reveal (1993)
Empetrales Raf. (1838)
Rhodorales Horan. (1847)
Vacciniales Dumort. (1829)
1. Actinidiaceae Hutch., nom. cons. (1926)
Saurauiaceae J. Agardh, nom. cons. (1858)
2. Clethraceae Klotzsch, nom. cons. (1851)
3. Cyrillaceae Endl. (1841)
4. Ericaceae Juss., nom. cons. (1789)
Andromedaceae (Endl.)DC. ex Schnizl. (1843-1870)
Arbutaceae (Meisn.) Bromhead (1840)
Arctostaphylaceae J. Agardh (1858)
Azaleaceae Vest (1818)
Diplarchaceae Klotzsch (1860)
Kalmiaceae Durande (1782)
Ledaceae Link (1821)
Menziesiaceae Klotzsch (1851)
Rhododendraceae Juss. (1789)
Rhodoraceae Vent. (1799)
Salaxidaceae J. Agardh (1858)
Siphonandraceae Klotzsch, nom. illeg. (1851)
Vacciniaceae DC. ex Gray, nom. cons. (1821)
5. Empetraceae Gray, nom. cons. (1821)
6. Monotropaceae Nutt., nom. cons. (1818)
Hypopityaceae Link (1831)
7. Pyrolaceae Dumort., nom. cons. (1829)
8. Epacridaceae R. Br. (1810)
Prionotaceae Hutch. (1969)
Stypheliaceae Horan. (1834)
4. Stylidiales Takht. ex Reveal (1992)
1. Stylidiaceae R. Br., nom. cons. (1810)
Candolleaceae F. Muell., nom. illeg. (1883)
Donatiaceae (Engl.) B. Chandler, nom. cons. (1911)
5. Sarraceniales Bromhead (1838)
1. Sarraceniaceae Dumort., nom. cons. (1829)
Heliamphoraceae Chrtek, Slavíková & Studnicka (1992)
22. Cornanae Thorne ex Reveal (1996)
Dipsacanae Takht. (1997)
Eucommianae Takht. ex Reveal (1993)
1. Cornales Dumort. (1829)
Adoxales Nakai (1949)
Aquifoliales Senft (1856)
Aralidiales Takht. ex Reveal (1992)
Aucubales Takht. (1997)
Garryales Lindl. (1846)
Griseliniales Takht. (1997)
Helwingiales Takht. (1997)
Hortensiales Griseb. (1854)
Hydrangeales Nakai (1943)
Icacinales Tiegh. ex Reveal (1993)
Metteniusales Takht. (1997)
Roridulales Nakai (1943)
Torricelliales Takht. ex Reveal (1992)
Viburnales Dumort. (1829)
1. Garryaceae Lindl., nom. cons. (1834)
2. Alangiaceae DC., nom. cons. (1828)
Metteniusaceae H. Karst. ex Schnizl. (1843-1870)
3. Nyssaceae Juss. ex Dumort., nom. co˙ns. (1829)
4. Cornaceae (Bercht. & J. Presl) Dumort., nom. cons. (1829)
Corokiaceae Kapil ex Takht. (1997)
Curtisiaceae (Harms) Takht. (1987)
Griseliniaceae J.R. Forst. & G. Forst. ex A. Cunn. (1839)
Kaliphoraceae Takht. (1996)
Mastixiaceae Calest. (1905)
Melanophyllaceae Takht. ex Airy Shaw (1972)
5. Roridulaceae Engl. & Gilg, nom. cons. (1924)
6. Davidiaceae (Harms) H.L. Li (1955)
7. Escalloniaceae R. Br. ex Dumort., nom. cons. (1829)
Abrophyllaceae Nakai (1943)
Argophyllaceae (Engl.) Takht. (1987)
Carpodetaceae Fenzl (1841)
Polyosmaceae Blume (1851)
8. Helwingiaceae Decne. (1836)
9. Torricelliaceae (Wang.) Hu (1934)
10. Aucubaceae J. Agardh (1858)
11. Aralidiaceae Philipson & B.C. Stone (1980)
12. Diapensiaceae (Link) Lindl., nom. cons. (1836)
Galacaceae D. Don (1828)
13. Phellinaceae (Loes.) Takht. (1967)
14. Aquifoliaceae Bartl., nom. cons. (1830)
Ilicaceae Bercht. & J. Presl (1820)
15. Paracryphiaceae Airy Shaw (1964)
16. Sphenostemonaceae P. Royen & Airy Shaw (1972)
17. Symplocaceae Desf., nom. cons. (1820)
18. Icacinaceae (Benth.) Miers, nom. cons. (1851)
Pennantiaceae J. Agardh (1858)
Phytocrenaceae Arn. ex R. Br. (1852)
19. Montiniaceae Nakai (1943)
20. Columelliaceae D. Don, nom. cons. (1828)
21. Alseuosmiaceae Airy Shaw (1964)
22. Hydrangeaceae Dumort., nom. cons. (1829)
Hortensiaceae Bercht. & J. Presl (1820)
Kirengeshomaceae Nakai (1943)
Philadelphaceae Martinov (1820)
23. Sambucaceae Batsch (1786)
24. Viburnaceae Raf. (1820)
Tinaceae Martinov (1820)
25. Menyanthaceae (Dumort.) Dumort., nom. cons. (1829)
26. Adoxaceae Trautv., nom. cons. (1853)
27. Phyllonomaceae Small (1905)
Dulongiaceae J. Agardh, nom. illeg. (1858)
28. Tribelaceae (Engl.) Airy Shaw (1964)
29. Eremosynaceae Dandy (1959)
30. Pterostemonaceae Small, nom. cons. (1905)
31. Tetracarpaeaceae Nakai (1943)
2. Eucommiales Nemejc ex Cronquist (1981)
1. Eucommiaceae Engl., nom. cons. (1909)
3. Dipsacales Dumort. (1829)
Caprifoliales Lindl. (1833)
Lonicerales T. Liebe (1866)
1. Caprifoliaceae Juss., nom. cons. (1789)
Carlemanniaceae Airy Shaw (1965)
Loniceraceae Vest (1818)
2. Valerianaceae Batsch, nom. cons. (1802)
Triplostegiaceae (Höck) Airy Shaw (1964)
3. Dipsacaceae Juss., nom. cons. (1789)
Scabiosaceae Adans. (1763)
4. Morinaceae Raf. (1820)
5. Calyceraceae R. Br. ex Rich., nom. cons. (1820)
Boopidaceae Cass. (1816)
23. Loasanae R. Dahlgren ex Reveal (1996)
1. Loasales Bessey (1907)
1. Loasaceae Dumort., nom. cons. (1822)
Cevalliaceae Griseb. (1854)
Gronoviaceae Endl. (1841)
24. Gentiananae Thorne ex Reveal (1992)
Oleanae Takht. (1997)
1. Goodeniales Lindl. (1833)
Brunoniales Lindl. (1833)
1. Goodeniaceae R. Br. (1810)
Brunoniaceae Dumort., nom. cons. (1829)
Scaevolaceae Lindl. (1830)
2. Oleales Lindl. (1833)
Jasminales Dumort. (1829)
1. Oleaceae Hoffmanns. & Link, nom. cons. (1813-1820)
Bolivariaceae Griseb. (1838)
Forestieraceae Endl. (1841)
Fraxinaceae Vest (1818)
Jasminaceae Adans. (1763)
Ligustraceae G. Mey. (1836)
Lilacaceae Vent., nom. illeg. (1799)
Schreberaceae (Wight) Schnizl. (1843-1870)
Syringaceae Horan. (1847)
3. Gentianales Lindl. (1833)
Apocynales Bromhead (1838)
Asclepiadales Dumort. (1829)
Chironiales Griseb. (1854)
Cinchonales Lindl. (1835)
Desfontainiales Takht. (1997)
Galiales Bromhead (1838)
Loganiales Lindl. (1833)
Rubiales Dumort. (1829)
Theligonales Nakai (1942)
Vincales Horan. (1847)
1. Desfontainiaceae Endl., nom. cons. (1841)
2. Loganiaceae R. Br. ex Mart., nom. cons. (1827)
Antoniaceae Hutch. (1959)
Gardneriaceae Wall. ex Perleb (1838)
Gelsemiaceae (G. Don) Struwe & V. Albert (1995)
Geniostomaceae Struwe & V. Albert (1995)
Plocospermataceae Hutch. (1973)
Potaliaceae Mart. (1827)
Spigeliaceae Mart. (1827)
Strychnaceae DC. ex Perleb (1818)
3. Dialypetalanthaceae Rizzini & Occhioni, nom. cons. (1948)
4. Rubiaceae Juss., nom. cons. (1789)
Aparinaceae Adans. (1763)
Asperulaceae Spenn. (1835)
Catesbaeaceae Martinov (1820)
Cephalanthaceae Raf. (1820)
Cinchonaceae Batsch (1802)
Coffeaceae Batsch (1802)
Coutareaceae Martinov (1820)
Galiaceae Lindl. (1836)
Gardeniaceae Dumort. (1829)
Guettardaceae Batsch (1802)
Hedyotidaceae Dumort. (1822)
Henriqueziaceae Bremek. (1957)
Houstoniaceae Raf. (1840)
Hydrophylacaceae Martinov (1820)
Lippayaceae Meisn. (1838)
Lygodisodeaceae Bartl. (1830)
Naucleaceae (DC.) Wernh. (1911)
Nonateliaceae Martinov (1820)
Operculariaceae Juss. ex Perleb (1818)
Pagamaeaceae Martinov (1820)
Psychotriaceae Rudolphi (1830)
Randicaceae Martinov (1820)
Sabiceaceae Martinov (1820)
5. Theligonaceae Dumort., nom. cons. (1829)
Cynocrambaceae Endl., nom. illeg. (1841)
6. Gentianaceae Juss., nom. cons. (1789)
Chironiaceae Horan. (1843)
Coutoubeaceae Martinov (1820)
Obolariaceae Martinov (1820)
7. Saccifoliaceae Maguire & Pires (1978)
8. Apocynaceae Juss., nom. cons. (1789)
Cerberaceae Martinov (1820)
Ophioxylaceae Mart. ex Perleb (1838)
Pacouriaceae Martinov (1820)
Plumeriaceae Horan. (1834)
Vincaceae Vest (1818)
Willughbeaceae J. Agardh (1858)
9. Asclepiadaceae R. Br., nom. cons. (1810)
Cynanchaceae G. Mey. (1836)
Periplocaceae Schltr., nom. cons. (1905)
Stapeliaceae Horan. (1834)
25. Lamianae Takht. (1967)
1. Lamiales Bromhead (1838)
Acanthales Lindl. (1833)
Bignoniales Lindl. (1833)
Callitrichales Dumort. (1829)
Gesneriales Dumort. (1829)
Globulariales Dumort. (1829)
Lentibulariales Lindl. (1833)
Pinguiculales Dumort. (1829)
Plantaginales Lindl. (1833)
Rhinanthales Dumort. (1829)
Scrophulariales Lindl. (1833)
Verbenales Horan. (1847)
1. Retziaceae Bartl. (1830)
2. Stilbaceae Kunth, nom. cons. (1831)
3. Buddlejaceae K. Wilh. (1910)
4. Scrophulariaceae Juss., nom. cons. (1789)
Aeginetiaceae Livera (1927)
Antirrhinaceae Pers. (1807)
Aragoaceae D. Don (1835)
Buchneraceae (Benth.) Lilja (1870)
Caprariaceae Martinov (1820)
Chelonaceae Martinov (1820)
Digitalidaceae Martinov (1820)
Ellisiophyllaceae Honda (1930)
Erinaceae Duvau ex Pfeiff. (1873)
Euphrasiaceae Martinov (1820)
Gratiolaceae Martinov (1820)
Hebenstretiaceae Horan. (1834)
Limosellaceae J. Agardh (1858)
Linariaceae Martinov (1820)
Melampyraceae Rich. ex Hook. & Lindl. (1821)
Oftiaceae Takht. & Reveal (1993)
Orobanchaceae Vent., nom. cons. (1799)
Oxycladaceae (Miers) Schnizl. (1843-1870)
Paulowniaceae Nakai (1949)
Pedicularidaceae Juss. (1789)
Phelypaeaceae Horan. (1834)
Rhinanthaceae Vent. (1799)
Schlegeliaceae (A.H. Gentry) Reveal (1996)
Selaginaceae Choisy, nom. cons. (1823)
Sibthorpiaceae D. Don (1835)
Spielmanniaceae J. Agardh, nom. illeg. (1858)
Verbasaceae Raf. (1821)
Veronicaceae Durande (1782)
5. Myoporaceae R. Br., nom. cons. (1810)
Bontiaceae Horan. (1834)
6. Globulariaceae DC., nom. cons. (1805)
7. Plantaginaceae Juss., nom. cons. (1789)
Littorellaceae Gray (1821)
Psylliaceae Horan. (1834)
8. Lentibulariaceae Rich., nom. cons. (1808)
Pinguiculaceae Dumort. (1829)
Utriculariaceae Hoffmanns. & Link, nom. cons. (1813-1820)
9. Pedaliaceae R. Br., nom. cons. (1810)
Sesamaceae R. Br. ex Bercht. & J. Presl (1820)
10. Trapellaceae Honda & Sakisaka (1930)
11. Martyniaceae Stapf, nom. cons. (1895)
12. Gesneriaceae Dumort., nom. cons. (1822)
Belloniaceae Martinov (1820)
Besleriaceae Raf. (1838)
Cyrtandraceae Jack (1823)
Didymocarpaceae D. Don (1822)
Ramondaceae Godr. (1850)
13. Bignoniaceae Juss., nom. cons. (1789)
Crescentiaceae Dumort. (1829)
14. Acanthaceae Juss., nom. cons. (1789)
Justiciaceae Raf. (1838)
Mendonciaceae Bremek. (1954)
Meyeniaceae Sreem. (1977)
Nelsoniaceae (Nees) Sreem. (1977)
Thomandersiaceae Sreem. (1977)
Thunbergiaceae (Dumort.) Lilja (1870)
15. Verbenaceae J. St.-Hil., nom. cons. (1805)
Aegiphilaceae Raf. (1838)
Avicenniaceae Endl. (1841)
Chloanthaceae Hutch. (1959)
Cyclocheilaceae Marais (1981)
Durantaceae J. Agardh (1858)
Lantanaceae Martinov (1820)
Nesogenaceae Marais (1981)
Nyctanthaceae J. Agardh (1858)
Petreaceae J. Agardh (1858)
Phrymaceae Schauer, nom. cons. (1847)
Siphonanthaceae Raf. (1838)
Symphoremataceae (Meisn.) Mold. ex Reveal & Hoogland (1991)
Viticaceae Juss. (1789)
16. Lamiaceae Lindl., nom. cons. (1836)
Glechomaceae Martinov (1820)
Labiatae Juss., nom. cons. et nom. alt. (1789)
Melittidaceae Martinov (1820)
Menthaceae Burnett (1835)
Nepetaceae Horan. (1834)
Salazariaceae F.A. Barkley (1975)
Salviaceae Raf. (1837)
Scutellaraceae Caruel (1894)
Tetrachondraceae Wettst. (1924)
17. Callitrichaceae Link, nom. cons. (1821)
Stellariaceae MacMill., nom. illeg. (1892)
2. Hydrostachyales Diels ex Reveal (1993)
1. Hydrostachyaceae Engl., nom. cons. (1898)
3. Hippuridales Pulle ex Reveal (1996)
1. Hippuridaceae Link, nom. cons. (1821)
Magnolianae through Theanae, Rosanae through Aralianae, or Alismatanae through Pandananae
Next Series of Lecture Notes:
Reveal System of Angiosperm Classification
Lecture Schedule
15 Nov 1997
